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In a world, where peace is protected by a happy melody of "Legendary Partiture", sung by a fairy of Songs, Hammy,
the entity which brings about the melody of sorrow - King of Evil, Mephisto - has appeared.
The queen of Happiness, Aphrodite, scatters the notes of Legendary Partiture to protect the peace.
The queen entrusts a mission to find those who bear a special "mark" of being able to become a Pretty Cure to the fairy Hammy and seven fairy tones.
Together, they are dispatched to gather the notes and ressurect the partiture.
Meet Kitajou Hibiki, a middle school sophomore.
After she had a fight with her childhood friend, Minamino Kanade, she met a mysterious lady.
However, the lady was a subordinate of Mephisto!
The fairy of Songs, Hammy, rushes to troubled Hibiki, having found the "mark" in her heart.
Kanade and Hibiki, so used to quarrel with each other, unite their hearts and transform into Pretty Cure: "We can't let you do that anymore!"